1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China;
2. CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China;
3. State Key Laboratory of Palaeobiology and Stratigraphy (Nanjing Institute of Geology and Palaeontology, CAS), Nanjing 210008, China;
4. University of Chinese Academy of Sciences, Beijing 100049, China